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Biological Evolution

Code: 60222
ECTS: 6.0
Lecturers in charge: prof. dr. sc. Damjan Franjević - Lectures
Lecturers: prof. dr. sc. Damjan Franjević - Seminar

Josip Skejo, mag. biol. exp. - Practicum
Take exam: Studomat
English level:


All teaching activities will be held in Croatian. However, foreign students in mixed groups will have the opportunity to attend additional office hours with the lecturer and teaching assistants in English to help master the course materials. Additionally, the lecturer will refer foreign students to the corresponding literature in English, as well as give them the possibility of taking the associated exams in English.

1. komponenta

Lecture typeTotal
Lectures 30
Practicum 30
Seminar 15
* Load is given in academic hour (1 academic hour = 45 minutes)
1. Recognising the evolutionary issues in the biological science
2. Know the basic mechanisms of evolution in biology
3. Connect the acquired knowledge and develop critical thinking in understanding the evolutionary mechanisms and theory
4. Explain the evolutionary mechanisms of acquired knowledge
5. Illustrate evolutionary mechanisms in biological science
6. Use the phylogenetic analyses in the evolution
7. Apply mathematical models in phylogeny and evolution
8. Motivate students to collect new literature data for the purpose of interpreting phylogenetic relationships and evolutionary science

Subject integrates content from lectures, practical exercises and seminar papers:
1. Evolution as a scientific discipline and its historical development
2. Material evidences of evolution (paleontology, comparative anatomy, molecular biology, genetics, physiology, biogeography)
3. Massive extinctions and the rate of evolutionary processes
4. Origin of life (chemical and biological evolution)
5. Evolution of the eukaryotic cell, endosymbiosis (organelle evolution)
6. Evolution of the Metazoa
7. Levels of evolutionary processes (microevolution, macroevolution, megaevolution)
8. Speciation
9. Isolation mechanisms
10. Factors of evolution (variability, mutation, recombination, migration, selection, genetic drift)
11. Epigenetics and Darwinism
12. Molecular evolution
13. Molecular phylogenetics
14. Population genetics and evolution
15. Human evolution

1., 2. Cosmic evolution
3. Origin of life
4., 5. The evolution of eukaryotic cell
6. Material evidences of evolution
7., 14. Molecular evolution and phylogeny
15. Population genetics

Seminars are linked to each lecture:
1. Evolution of the Universe (Higgs boson "God particle")
2. The origin of life theory
3. Definitions of living systems
4. Universal ancestors of living beings
5. Evolution of eukaryotic cell (endosymbiosis)
6. Evolution of metazoans
7. Phylogenetic trees
8. The rate of evolutionary processes
9. Fossils (ancient DNA)
10. Megaevolution, macroevolution, philostratigraphy
11. Molecular mechanisms of variability
12. Selection
13. Mimicry and convergence
14. Human Evolution
15. Theories of migrations from Africa and the colonization of continents
  1. Stephen C. Stearns, Rolf F. Hoekstra: Evolution an introduction 2nd Edition, Oxford University Press, 2005.
  2. Wen-Hsiung Li: Molecular evolution, Sinauer Assotiates, Inc., 1997.
  3. Douglas J. Futuyma: Evolutionary Biology, Sinauer Assotiates, Inc., 1997.
  4. Peter W. Price: Biological evolution, Saunders College Publishing, 1996.
  5. Marcello Barbieri: The Organic Codes, Cambridge University Press, 2003.
  6. Mirjana Kalafatić: Osnove biološke evolucije, Priroda, Zagreb, 1998.
  7. Roger Lewin: Human Evolution An Illustrated Introduction, Blackwell Publishing Ltd, 2005.
  8. Libor Ebringer, Juraj Krajčovič: Cell origin and evolution, VEDA, Bratislava, 1994.
  9. Barry G. Hall: Phlogenetic Trees Made Easy, 4th Edition, Sinauer Associates, Inc. Publishers, 2011.
Prerequisit for:
Enrollment :
Passed : Genetics
5. semester
Mandatory course - Regular study - Biology
Consultations schedule: